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The effects of action-based predictions in early visual cortex.

Code ↔ Paper

4 matches between paragraphs of the paper and lines of its authors' code, computed by the harvester (lexical-v1). Click a colored paragraph or line to see its counterpart.

The 4 matches
  1. [1] § STAR★Methods › Method details › Experimental design ↔ Code.zip/Behavioural/actPredAllTraining_v2.py, lines 125–171 · score 0.73 · wrong button, rightward gratings, passive block, clockwise, leftward, soon
  2. [2] § STAR★Methods › Quantification and statistical analysis › Eye tracking analysis ↔ Code.zip/fMRI/actPred_v2.py, lines 90–115 · score 0.61 · SR Research, EyeLink, calibration, eyetracker, tracking
  3. [3] § STAR★Methods › Method details › Experimental design ↔ Code.zip/Behavioural/actPredGratingTraining_v2.py, lines 93–117 · score 0.59 · rightward gratings, fixation dot, clockwise, leftward, counterbalanced, trained
  4. [4] § STAR★Methods › Quantification and statistical analysis › MRI analysis ↔ Code.zip/Behavioural/actPredGratingTraining_v2.py, lines 93–117 · score 0.54 · rightward gratings, fixation dot, leftward, space, predictors, cue

Paper

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The authors' code

Python · 363 lines · 13 KB · CC-BY-4.0 · 2 matches

  1. #!/usr/bin/env python3
  2. # -*- coding: utf-8 -*-
  3. """
  4. Created on Wed Jul 7 16:13:47 2021
  5. @author: bianca
  6. """
  7. ##############################################################################
  8. #
  9. # ActPred3T_WP1
  10. #
  11. # !! Updated version !!
  12. # Cue is now fixation dot changing colour instead of enlarging
  13. #
  14. # Script to train mapping between tone and gratings.
  15. #
  16. # BvK 07-07-2021
  17. ##############################################################################
  18. from psychopy import prefs
  19. prefs.hardware['audioLib'] = ['PTB']
  20. from psychopy import visual, core, event, sound, data, gui
  21. import os, datetime, random, pickle
  22. import numpy as np
  23. # Add escape and q as escape keys
  24. #for key in ['q', 'escape']:
  25. # event.globalKeys.add(key, func=core.quit)
  26. # Paths
  27. #root_dir = '/analyse/Project0281/ActPred3T/'
  28. #src_dir = '/analyse/Project0281/ActPred3T/Behavioural/'
  29. #root_dir = 'C:/Users/Bianca/Work/Experiments/Glasgow_3T/Code_exp/'
  30. #src_dir = 'C:/Users/Bianca/Work/Experiments/Glasgow_3T/Code_exp/Behavioural/'
  31. root_dir = '/home/bianca/Work/Experiments/Glasgow_3T/Code_exp/actpred3T/' #linux laptop
  32. src_dir = root_dir + 'Behavioural/'
  33. #RT_dir = '/analyse/Project0281/ActPred3T/Behavioural/'
  34. RT_dir = root_dir + 'Behavioural/'
  35. # GUI to input participant number etc
  36. expName = 'actpred3T_gratingtraining'
  37. expInfo = {'Date':data.getDateStr(), 'Run':['1','2','3','4','5'], 'Participant_number': '', 'Participant_ID':''}
  38. diabox = gui.DlgFromDict(dictionary=expInfo,
  39. title=expName,
  40. order=['Participant_number', 'Participant_ID', 'Run', 'Date'],
  41. fixed=['Date'])
  42. s = int(expInfo['Participant_number']);
  43. run = int(expInfo['Run']);
  44. date = expInfo['Date']
  45. print('expinfo:', expInfo['Participant_number'])
  46. print('s = ', s)
  47. saving_dir = src_dir + 'data/p' + str(s) +'/'
  48. if not os.path.exists(saving_dir):
  49. os.makedirs(saving_dir)
  50. logFile = open(saving_dir + 'gratingtraining_p' + str(s) + '_run' + str(run) +'_' + date + '.txt','w')
  51. logFile.write('Date: \t' + expInfo['Date'] + '\n')
  52. logFile.write('Participant number: \t' + expInfo['Participant_number'] + '\n')
  53. logFile.write('Participant_ID: \t' + expInfo['Participant_ID'] + '\n')
  54. logFile.write('Run: \t' + expInfo['Run'] + '\n')
  55. logFile.write('Start: \t' + str(datetime.datetime.now()) + '\n')
  56. logFile.write('trialN\tPart\tCond\tPredCueOn\tpredCueOff\tCueOn\tCueOff\tTActKeyPressed\tActKeyPressed\tRT\tOrient\tStartStim1\tStopStim1\tITI\n')
  57. # Window
  58. #win = visual.Window(monitor='Laptop', units = 'deg', fullscr = True)
  59. #win = visual.Window([600,600], monitor='Laptop', units = 'deg')
  60. win = visual.Window([600,600], monitor='HomeMon', units = 'deg')
  61. #win = visual.Window([800,800], monitor='WorkPC', units = 'deg')
  62. #win = visual.Window(monitor='WorkPC', units = 'deg', fullscr = True)
  63. win.mouseVisible = False
  64. circle_color_space = 'rgb'
  65. subList = list(range(1,25)) #1:24
  66. # Load pickle file
  67. subsMaps = pickle.load(open(root_dir + 'mapping', 'rb'))
  68. # Randomise trial order for each run
  69. seed = random.sample(range(1000), k=1)
  70. random.seed(seed[0])
  71. conds_act = np.array([1,1,1,1,1,2,2,2,2,2])
  72. random.shuffle(conds_act)
  73. seed = random.sample(range(1000), k=1)
  74. random.seed(seed[0])
  75. conds_pas = np.array([4,4,4,4,4,5,5,5,5,5])
  76. random.shuffle(conds_pas)
  77. # Define all screens/stimuli
  78. part1aScreen = visual.TextStim(win, '''Part 1 \n\nWhen you hear a high tone, you will see a leftward grating. When you hear a low tone, you will see a rightward grating. \nPay close attention to the stimuli and learn which tone predicts which grating.
  79. \n\nPress space bar to continue''', wrapWidth=20)
  80. part1bScreen = visual.TextStim(win, '''Part 1 \n\nWhen you hear a high tone, you will see a rightward grating. When you hear a low tone, you will see a leftward grating. \nPay close attention to the stimuli and learn which tone predicts which grating.
  81. \n\nPress space bar to continue''', wrapWidth=20)
  82. part2aScreen = visual.TextStim(win, '''Part 2 \n\nWhen you hear a high tone, you will see a leftward grating. When you hear a low tone, you will see a rightward grating. \nPay close attention to the stimuli and learn which tone predicts which grating.
  83. \n\nPress space bar to continue''', wrapWidth=20)
  84. part2bScreen = visual.TextStim(win, '''Part 2 \n\nWhen you hear a high tone, you will see a rightward grating. When you hear a low tone, you will see a leftward grating. \nPay close attention to the stimuli and learn which tone predicts which grating.
  85. \n\nPress space bar to continue''', wrapWidth=20)
  86. part2Screen = visual.TextStim(win, '''Part 2 \n\nWhen you hear the tone, prepare for the corresponding grating to be presented. It will be presented after the fixation dot enlarges.
  87. \n\nPress space bar to continue''', wrapWidth=20)
  88. breakScreen = visual.TextStim(win, 'Questions?')
  89. predCueLow = sound.Sound(400,volume = 0.846, secs = 0.3)
  90. predCueHigh = sound.Sound(1000,volume = 0.8, secs = 0.3)
  91. predCueNo = sound.Sound(100,volume = 0.8, secs = 0.3)
  92. fix = visual.Circle(win, radius=0.15, units='deg', edges=100, lineColorSpace=circle_color_space, lineColor=[-1,-1,-1], fillColor=[-1,-1,-1], fillColorSpace=circle_color_space)
  93. cue = visual.Circle(win, radius=0.15, units='deg', edges=100, lineColorSpace=circle_color_space, lineColor=[1,-1,-1], fillColor=[1,-1,-1], fillColorSpace=circle_color_space)
  94. refStimLeft = visual.GratingStim(win,tex='sin', mask='raisedCos', maskParams = {'fringeWidth':0.1},contrast = 0.5, units='deg', ori = -45, size = 8, sf = 1.5)
  95. testStimLeft = visual.GratingStim(win,tex='sin', mask='raisedCos', maskParams = {'fringeWidth':0.1},contrast = 0.5, units='deg', size = 8, sf = 1.5)
  96. refStimRight = visual.GratingStim(win,tex='sin', mask='raisedCos', maskParams = {'fringeWidth':0.1},contrast = 0.5, units='deg', ori = 45, size = 8, sf = 1.5)
  97. testStimRight = visual.GratingStim(win,tex='sin', mask='raisedCos', maskParams = {'fringeWidth':0.1},contrast = 0.5, units='deg', size = 8, sf = 1.5)
  98. annulus = visual.Circle(win, radius=0.25, units='deg', edges=100, lineWidth = 0, fillColor = [0,0,0])
  99. refSound = sound.Sound(400, volume = 0.8, secs = 0.5)
  100. quest = visual.TextStim(win, 'Clockwise or counterclockwise?', wrapWidth=20)
  101. endScreen = visual.TextStim(win, 'End of run')
  102. def trial_sequence(cue_type, button, orient):
  103. pred_cue(cue_type)
  104. pas_trial()
  105. present_grating(orient)
  106. ITI()
  107. def pred_cue(cue_type):
  108. fix.draw()
  109. win.flip()
  110. if cue_type == 'high':
  111. predCueHigh.play()
  112. elif cue_type == 'low':
  113. predCueLow.play()
  114. elif cue_type == 'no':
  115. predCueNo.play()
  116. predCueOn = globalClock.getTime()
  117. cueClock.reset()
  118. cueClock.add(0.3)
  119. while cueClock.getTime() < 0:
  120. pass
  121. predCueOff = globalClock.getTime()
  122. logFile.write('\t'.join([str(predCueOn), str(predCueOff)])+'\t')
  123. def present_grating(orient):
  124. stimClock.reset()
  125. stimClock.add(0.2)
  126. while stimClock.getTime() < 0:
  127. pass
  128. if orient == 'left':
  129. refStimLeft.draw()
  130. elif orient == 'right':
  131. refStimRight.draw()
  132. annulus.draw()
  133. fix.draw()
  134. win.flip()
  135. stim1On = globalClock.getTime()
  136. stimClock.reset()
  137. stimClock.add(0.5)
  138. while stimClock.getTime() < 0:
  139. pass
  140. stim1Off = globalClock.getTime()
  141. if orient == 'left':
  142. logFile.write('\t'.join([str(-45), str(stim1On), str(stim1Off)])+'\t')
  143. elif orient == 'right':
  144. logFile.write('\t'.join([str(45), str(stim1On), str(stim1Off)])+'\t')
  145. def pas_trial():
  146. fix.draw()
  147. win.flip()
  148. fixClock.reset()
  149. fixClock.add(3)
  150. while fixClock.getTime() < 0:
  151. pass
  152. cue.draw()
  153. win.flip()
  154. cueOn = globalClock.getTime()
  155. cueClock.reset()
  156. cueClock.add(0.3)
  157. while cueClock.getTime() < 0:
  158. pass
  159. cueOff = globalClock.getTime()
  160. fix.draw()
  161. win.flip()
  162. RTClock.reset()
  163. #RTClock.add(RT_act)
  164. RTClock.add(1-0.2)
  165. while RTClock.getTime() < 0:
  166. pass
  167. logFile.write('\t'.join([str(cueOn), str(cueOff), str(-1), str(0), str(-1)])+'\t')
  168. def ITI():
  169. fix.draw()
  170. win.flip()
  171. ITIClock.reset()
  172. ITIClock.add(1.5)
  173. while ITIClock.getTime() < 0:
  174. pass
  175. logFile.write('\t'.join([str(1.5)])+'\n')
  176. globalClock = core.Clock()
  177. blockClock = core.Clock()
  178. cueClock = core.Clock()
  179. RTClock = core.Clock()
  180. stimClock = core.Clock()
  181. fixClock = core.Clock()
  182. isiClock = core.Clock()
  183. questClock = core.Clock()
  184. ITIClock = core.Clock()
  185. exit = False
  186. if subsMaps[s] == 1 or subsMaps[s] == 3:
  187. part1aScreen.draw()
  188. elif subsMaps[s] == 2 or subsMaps[s] == 4:
  189. part1bScreen.draw()
  190. win.flip()
  191. keys = ['']
  192. while keys[0] not in ['escape', 'space']:
  193. keys = event.waitKeys()
  194. if keys[0] == 'space':
  195. pass
  196. elif keys[0] == 'escape':
  197. exit = True
  198. win.close()
  199. core.quit()
  200. fix.draw()
  201. win.flip()
  202. blockClock.reset()
  203. blockClock.add(3)
  204. while blockClock.getTime() < 0:
  205. pass
  206. counterL = 0
  207. counterR = 0
  208. counterAct = 0
  209. counterPas = 0
  210. for trial, cond in enumerate(conds_pas,1):
  211. part = 1
  212. logFile.write('\t'.join([str(trial), str(part), str(cond)])+'\t')
  213. if cond == 4:
  214. if subsMaps[s] == 1:
  215. trial_sequence('high', 'left', 'left')
  216. counterL = counterL + 1
  217. counterPas = counterPas + 1
  218. elif subsMaps[s] == 2:
  219. trial_sequence('low', 'right', 'left')
  220. counterL = counterL + 1
  221. counterPas = counterPas + 1
  222. elif subsMaps[s] == 3:
  223. trial_sequence('high', 'right', 'left')
  224. counterL = counterL + 1
  225. counterPas = counterPas + 1
  226. elif subsMaps[s] == 4:
  227. trial_sequence('low', 'left', 'left')
  228. counterL = counterL + 1
  229. counterPas = counterPas + 1
  230. elif cond == 5:
  231. if subsMaps[s] == 1:
  232. trial_sequence('low', 'right', 'right')
  233. counterR = counterR + 1
  234. counterPas = counterPas + 1
  235. elif subsMaps[s] == 2:
  236. trial_sequence('high', 'left', 'right')
  237. counterR = counterR + 1
  238. counterPas = counterPas + 1
  239. elif subsMaps[s] == 3:
  240. trial_sequence('low', 'left', 'right')
  241. counterR = counterR + 1
  242. counterPas = counterPas + 1
  243. elif subsMaps[s] == 4:
  244. trial_sequence('high', 'right', 'right')
  245. counterR = counterR + 1
  246. counterPas = counterPas + 1
  247. #Break
  248. breakScreen.draw()
  249. win.flip()
  250. keys = event.waitKeys()
  251. #Prepare part2
  252. if subsMaps[s] == 1 or subsMaps[s] == 3:
  253. part2aScreen.draw()
  254. elif subsMaps[s] == 2 or subsMaps[s] == 4:
  255. part2bScreen.draw()
  256. win.flip()
  257. keys = ['not pressed']
  258. while keys[0] not in ['escape', 'space']:
  259. keys = event.waitKeys()
  260. if keys[0] == 'space':
  261. pass
  262. elif keys[0] == 'escape':
  263. exit = True
  264. win.close()
  265. core.quit()
  266. fix.draw()
  267. win.flip()
  268. blockClock.reset()
  269. blockClock.add(3)
  270. while blockClock.getTime() < 0:
  271. pass
  272. counterL = 0
  273. counterR = 0
  274. counterAct = 0
  275. counterPas = 0
  276. for trial, cond in enumerate(conds_pas,1):
  277. part = 2
  278. logFile.write('\t'.join([str(trial), str(part), str(cond)])+'\t')
  279. if cond == 4:
  280. if subsMaps[s] == 1:
  281. trial_sequence('high', 'left', 'left')
  282. counterL = counterL + 1
  283. counterPas = counterPas + 1
  284. elif subsMaps[s] == 2:
  285. trial_sequence('low', 'right', 'left')
  286. counterL = counterL + 1
  287. counterPas = counterPas + 1
  288. elif subsMaps[s] == 3:
  289. trial_sequence('high', 'right', 'left')
  290. counterL = counterL + 1
  291. counterPas = counterPas + 1
  292. elif subsMaps[s] == 4:
  293. trial_sequence('low', 'left', 'left')
  294. counterL = counterL + 1
  295. counterPas = counterPas + 1
  296. elif cond == 5:
  297. if subsMaps[s] == 1:
  298. trial_sequence('low', 'right', 'right')
  299. counterR = counterR + 1
  300. counterPas = counterPas + 1
  301. elif subsMaps[s] == 2:
  302. trial_sequence('high', 'left', 'right')
  303. counterR = counterR + 1
  304. counterPas = counterPas + 1
  305. elif subsMaps[s] == 3:
  306. trial_sequence('low', 'left', 'right')
  307. counterR = counterR + 1
  308. counterPas = counterPas + 1
  309. elif subsMaps[s] == 4:
  310. trial_sequence('high', 'right', 'right')
  311. counterR = counterR + 1
  312. counterPas = counterPas + 1
  313. endScreen.draw()
  314. win.flip()
  315. event.waitKeys(keyList=['escape'])
  316. win.close()
  317. core.quit()

actPredGratingTraining_v2.py, under CC-BY-4.0 · at the source

Overview

Authors: Bianca M. van Kemenade1,2, Lars F. Muckli2
  1. Center for Psychiatry, Justus Liebig University Giessen, Klinikstrasse 36, 35392 Giessen, Germany
  2. Centre for Cognitive Neuroimaging (CCNi), School of Psychology and Neuroscience, University of Glasgow, 62 Hillhead Street, Glasgow, UK
Institutions: Justus-Liebig-Universität Gießen (Germany); University of Glasgow (United Kingdom)
Journal: iScience, volume 29, issue 9, article 117074
Dates: received 3 October 2025; accepted 20 July 2026; published online 18 August 2026
Type: Research article · Language: English
License: CC BY
Identifiers: DOI 10.1016/j.isci.2026.117074 · PMID 42662421 · PMCID PMC13520625 · OpenAlex W4413900552
Open access: gold, a free copy (OpenAlex)
Status: code verified
Categories: fMRI (modality), systems (subfield)
Methods: Statistics, Machine learning, Preprocessing, fMRI & imaging, Physiology & signal measures
Keywords: voluntary action, visual perception, early visual cortex, fMRI
Topic: Neural dynamics and brain function (Cognitive Neuroscience, Neuroscience), according to OpenAlex
Citations: not cited yet (Europe PMC); 55 references in the paper
Research resources: Matlab R2019b RRID:SCR_001622, PsychoPy 3.2.4 RRID:SCR_006571, LibSVM RRID:SCR_010243, BrainVoyager 22.0 for Linux RRID:SCR_013057

Abstract

Voluntary action typically results in reduced sensitivity to sensory action outcomes. The forward model theory proposes that this is due to neural suppression. However, this theory has recently been challenged by the pre-activation account, sharpening, and opposing process theory. In this fMRI study, we compared these theories by using univariate and multivariate analyses. Participants performed a visual orientation discrimination task on two sequential gratings, which were presented automatically (passive condition) or triggered by button press (active condition). Decoding of predicted stimulus orientation from early visual cortex activity prior to stimulus presentation was significantly above chance in both active and passive conditions. During stimulus presentation, actively generated stimuli elicited larger blood-oxygen-level-dependent (BOLD) responses. However, both decoding accuracy and the timing of the BOLD responses did not differ between conditions. These results differ from the forward model’s predicted sensory attenuation, the pre-activation account’s earlier BOLD response, and the enhanced precision of the sharpening hypothesis. Instead, they mostly fit the opposing process theory, which posits pre-activation in both conditions. However, the stronger BOLD response for actively generated stimuli is not predicted by any existing theory, implicating additional mechanisms, such as heightened attention or motor-related enhancement.

Reproduced under the paper's license (CC BY), from the paper cited above.

Repository

Its files are read in the Code ↔ Paper reader above, with 4 matches between paragraphs and lines of code.

Zenodo 20507627

License: CC-BY-4.0
State: the link answers, verified on 27 September 2026
Evidence: files inventoried
Size: 3 files
Software Heritage: not checked
Found in: “Data and code availability”
Not found: README, license file, CITATION.cff, environment file, tests, continuous integration, documentation
Tools: NumPy (8 files), PsychoPy (8 files)
Availability: 1 check, the latest on 27 September 2026: the link answers (HTTP 200)
  • 27 September 2026: the link answers (HTTP 200)
8 files

The paper's code and data availability statement is in the Data section.

Tracing map

Proposed by the machine: these links were found in the paper and verified at the source, without human review. The map will receive a Zenodo DOI once one of the paper's authors has validated it with their ORCID.

What the map holds:

  • 1 repository of the authors' code, each at its verified commit, with its license and how the link was found in the paper;
  • 8 scripts, each with its path and the digest of its content;
  • 4 matches between paragraphs of the paper and lines of the code (method lexical-v1);
  • neither the text of the paper nor the code itself.

Its JSON (tracing-map.json) is deposited on Zenodo with its DOI once the map is validated.

Data

No dataset and no data link were found in the paper.

Data and code availability

• Data have been deposited at Zenodo and are publicly available as of the date of publication at https://doi.org/10.5281/zenodo.20507627. • Code has been deposited at Zenodo and is publicly available as of the date of publication at https://doi.org/10.5281/zenodo.20507627. • For other inquiries, please contact the lead contact.

Reproduced under the paper's license (CC BY), from the paper cited above.

Versions

The history of this record: each version stored by the harvester or made by a correction of its authors or of the maintainers of its code, and what changed in its facts. The texts of the paper (its abstract, its availability statements) are not part of it; versions that changed only those are not listed.

Version 2, 28 September 2026

  • Authors: added Bianca M. van Kemenade (0000-0002-8631-9893); removed Bianca M. van Kemenade

Version 1, 27 September 2026: the first record

Recorded: type, language, journal, volume, issue, pages, dates, 2 authors, 4 keywords, 3 funders, 54 references, 4 RRIDs.

Cite

This paper

van Kemenade, B. M., & Muckli, L. F. (2026). The effects of action-based predictions in early visual cortex. iScience, 29(9), 117074. https://doi.org/10.1016/j.isci.2026.117074

BibTeX

@article{vankemenade2026effects,
author = {van Kemenade, Bianca M. and Muckli, Lars F.},
title = {{The effects of action-based predictions in early visual cortex}},
journal = {iScience},
year = {2026},
month = aug,
volume = {29},
number = {9},
pages = {117074},
publisher = {Elsevier},
issn = {2589-0042},
doi = {10.1016/j.isci.2026.117074},
url = {https://doi.org/10.1016/j.isci.2026.117074},
pmid = {42662421},
pmcid = {PMC13520625}
}

RIS

TY - JOUR
AU - van Kemenade, Bianca M.
AU - Muckli, Lars F.
TI - The effects of action-based predictions in early visual cortex
T2 - iScience
J2 - iScience
PY - 2026
DA - 2026/08/18
VL - 29
IS - 9
SP - 117074
SN - 2589-0042
PB - Elsevier
DO - 10.1016/j.isci.2026.117074
UR - https://doi.org/10.1016/j.isci.2026.117074
LA - en
ER -

CSL-JSON

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"title": "The effects of action-based predictions in early visual cortex",
"container-title": "iScience",
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"family": "van Kemenade",
"given": "Bianca M."
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{
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"given": "Lars F."
}
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"container-title-short": "iScience",
"volume": "29",
"issue": "9",
"page": "117074",
"DOI": "10.1016/j.isci.2026.117074",
"PMID": "42662421",
"PMCID": "PMC13520625",
"ISSN": "2589-0042",
"publisher": "Elsevier",
"URL": "https://doi.org/10.1016/j.isci.2026.117074",
"language": "en",
"issued": {
"date-parts": [
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18
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]
}
}

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Learning regularities in noise engages both neural predictive activity and representational changes.
Journal: Nature communications
In common: NumPy, 2 references
[6] doi:10.3758/s13428-026-03150-6 [code]
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Journal: Behavior research methods
In common: PsychoPy, NumPy, fMRI
[7] doi:10.1016/j.dcn.2026.101791 [code]
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Journal: Developmental cognitive neuroscience
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[8] doi:10.1162/nol.a.271 [code]
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Journal: Neurobiology of language (Cambridge, Mass.)
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[9] doi:10.1038/s41380-026-03694-1 [code]
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In common: PsychoPy, NumPy, fMRI
[10] doi:10.1038/s41467-026-72605-3 [code]
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Journal: Nature communications
In common: PsychoPy, NumPy, fMRI

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