OSCR

The human hippocampus can pattern separate memories by meaning.

Code ↔ Paper

9 matches between paragraphs of the paper and lines of its authors' code, computed by the harvester (lexical-v1). Click a colored paragraph or line to see its counterpart.

The 9 matches
  1. [1] § Behavioral Methods › Materials. › Covariate assessments. ↔ scripts/smst2_bids_participants_maker.py, lines 191–215 · score 0.70 · WAIS IV, Digit Symbol, subtests, Vocabulary, covariate, words
  2. [2] § Behavioral Methods › MRI Methods. › Hippocampal subfield segmentation. ↔ scripts/smst_mr1_HCmask_snapshot.sh, lines 1–50 · score 0.64 · coronal slices, template T1, brain, masks, SI
  3. [3] § Behavioral Methods › Materials. › Main task (sMST). › Stimuli. ↔ scripts/smst2_bids_tsv_maker.py, lines 303–386 · score 0.63 · Hungarian National Corpus, imageability, metric, arousal, concreteness, fillers
  4. [4] § Behavioral Methods › MRI Methods. › Analysis. ↔ scripts/smst_mr1_analysis.Rmd, lines 156–247 · score 0.61 · close lure, distant lure, lure correct, incorrect, hits, foil
  5. [5] § Results › RS Sensitive Clusters in the Hippocampal Head Differentially Activate for Exact Repeats and Modified Repeats During Encoding. ↔ scripts/smst_mr1_analysis.Rmd, lines 2190–2332 · score 0.61 · DG CA3, right hemisphere, left hemisphere, CA12, ROI, Facets
  6. [6] § Results › Neural Response Is Increased for Semantically Close Modified Phrases Compared to Exact Repetitions, with No Further Difference between Semantic Similarity Bins. ↔ scripts/smst_mr1_analysis.Rmd, lines 2190–2332 · score 0.54 · DG CA3, distant repeats, CA1, anatomically, ANOVA, head
  7. [7] § Behavioral Methods › Analysis. ↔ scripts/smst_mr1_analysis.Rmd, lines 398–440 · score 0.54 · Wald, confint, intercept, arousal, concreteness, GLMEMs
  8. [8] § Results › RS Sensitive Clusters in the Hippocampal Head Differentially Activate for Exact Repeats and Modified Repeats During Encoding. ↔ scripts/smst_mr1_analysis.Rmd, lines 442–540 · score 0.53 · Post hoc, covariate, clusters, encoding
  9. [9] § Results › Clusters Sensitive to Repetition at Encoding Do Not Track Mnemonic Discrimination Success within Lures but Differentiate Correctly Rejected Lures from Correctly Rejected Foils During Recogni ↔ scripts/smst_mr1_analysis.Rmd, lines 3552–3596 · score 0.50 · lure correct rejections, target hits, modified repeat, baseline, clusters

Paper

Loaded from Europe PMC by your browser, not stored by OSCR: doi.org · Europe PMC

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The authors' code

R Markdown · 5,188 lines · 155 KB · no license · 6 matches

The registry keeps no copy of this file: its repository has no license, so its authors keep all their rights to it. Your browser shows it from its source, with JavaScript.

It can be read at the source: scripts/smst_mr1_analysis.Rmd.

Overview

Authors: Alex Ilyés1,2,3, Borbála Mónika Brosig2,4, György Mező5,6,7, Attila Keresztes2,3,8
  1. Doctoral School of Psychology, Eötvös Loránd University, Budapest 1075, Hungary
  2. Brain Imaging Centre, Hungarian Research Network, Research Centre for Natural Sciences, Budapest 1117, Hungary
  3. Institute of Psychology, Eötvös Loránd University, Budapest 1064, Hungary
  4. Mental Health Sciences Division, Doctoral School, Semmelweis University, Budapest 1085, Hungary
  5. Konkoly Observatory, Hungarian Research Network, Research Centre for Astronomy and Earth Sciences, Budapest 1121, Hungary
  6. Wigner Data Center, Hungarian Research Network, Wigner Research Centre for Physics, Budapest 1121, Hungary
  7. Hungarian Academy of Sciences Centre of Excellence, Hungarian Research Network, Research Centre for Astronomy and Earth Sciences, Budapest 1121, Hungary
  8. Max Planck Partner Group of the Max Planck Institute for Human Development, Hippocampal Circuit and Code for Cognition Lab, Budapest 1117, Hungary
Dates: received 28 January 2026; accepted 18 June 2026; published online 28 July 2026; in print 4 August 2026
Type: Research article · Language: English
License: CC BY-NC-ND
Identifiers: DOI 10.1073/pnas.2603114123 · PMID 42520121 · PMCID PMC13438472 · OpenAlex W7171530764
Open access: hybrid, a free copy (OpenAlex)
Status: code verified
Categories: human (organism), cognitive (subfield)
Methods: Statistics, Preprocessing, fMRI & imaging
Keywords: pattern separation, episodic memory, semantic similarity, word2vec, repetition suppression
MeSH: Hippocampus*, Memory*, Adult, Brain Mapping, Female, Humans, Magnetic Resonance Imaging, Male, Memory, Episodic, Recognition, Psychology, Semantics, Young Adult (* major topic)
Journal subjects: Social Sciences, Psychological and Cognitive Sciences
Topic: Memory and Neural Mechanisms (Cognitive Neuroscience, Neuroscience), according to OpenAlex
Funding: Hungarian National Research, Development, and Innovation Office (FK128648 and FK146496); Max Planck Society (Max Planck Partner Group); Hungarian Academy of Sciences (0708-21 515 AT); Eotvos Lorand University (Doctoral Projects 2022 Doctoral Projects 2023); University Research Scholarship Program of the Ministry for Culture and Innovation (EKÖP-25-4-I-ELTE-1020, EKÖP-24-2-I-ELTE-687)
Citations: not cited yet (Europe PMC); 144 references in the paper

Abstract

The abstract is not reproduced here: the paper's license (CC BY-NC-ND) does not allow it. Read it in the paper, at the publisher or on Europe PMC.

Repositories

Its files are read in the Code ↔ Paper reader above, with 9 matches between paragraphs and lines of code.

Zenodo 18395407

License: CC-BY-4.0
State: the link answers, verified on 27 September 2026
Evidence: files inventoried
Size: 1 file
Software Heritage: not checked
Found in: the references
Not found: README, license file, CITATION.cff, environment file, tests, continuous integration, documentation
Tools: NumPy (6 files), PsychoPy (5 files), pandas (4 files)
Availability: 1 check, the latest on 27 September 2026: the link answers (HTTP 200)
  • 27 September 2026: the link answers (HTTP 200)
10 files
At the source:

Zenodo 20090797

License: CC-BY-4.0
State: the link answers, verified on 27 September 2026
Evidence: files inventoried
Size: 1 file
Software Heritage: not checked
Found in: the references
Not found: README, license file, CITATION.cff, environment file, tests, continuous integration, documentation
Tools: Curve Fitting Toolbox (2 files), afex (1 file), broom (1 file), easystats (1 file), emmeans (1 file), FSL (1 file), ggplot2 (1 file), ggpubr (1 file), lme4 (1 file), lmerTest (1 file), Matplotlib (1 file), NiBabel (1 file), NumPy (1 file), pandas (1 file), patchwork (1 file), psych (1 file), rstatix (1 file), SciPy (1 file), tidyverse (1 file)
Availability: 1 check, the latest on 27 September 2026: the link answers (HTTP 200)
  • 27 September 2026: the link answers (HTTP 200)
6 files
At the source:

ilyesalex/smst2_mr_experiment

License: none: the authors keep all their rights
State: the link answers, verified on 27 September 2026
Evidence: files inventoried
Commit: c73d841427fd9ac13d6434ad2bf9b971ab5c2e56, 27 January 2026
Languages: Python (9)
Size: 1,720 files, 9 scripts
Software Heritage: not archived
Found in: the Zenodo archive record
Holds: README
Not found: license file, CITATION.cff, environment file, tests, continuous integration, documentation
Tools: NumPy (6 files), PsychoPy (5 files), pandas (4 files)
Availability: 1 check, the latest on 27 September 2026: the link answers
  • 27 September 2026: the link answers
10 files, not copied: shown from their source

OSCR keeps no copy of these files: this repository has no license that allows it. The reader above shows each one from its source, fetched by your browser at commit c73d841, when its fingerprint is the one OSCR verified. How this works.

ilyesalex/smst2_mr_analysis

License: none: the authors keep all their rights
State: the link answers, verified on 27 September 2026
Evidence: files inventoried
Commit: b1f8280750fbe099b86224e616a30e5db4c0f85a, 8 May 2026
Languages: MATLAB (2), Shell (1), Python (1), R (1)
Size: 64 files, 5 scripts
Software Heritage: not archived
Found in: the Zenodo archive record
Holds: README, 1 notebook
Not found: license file, CITATION.cff, environment file, tests, continuous integration, documentation
Tools: Curve Fitting Toolbox (2 files), afex (1 file), broom (1 file), easystats (1 file), emmeans (1 file), FSL (1 file), ggplot2 (1 file), ggpubr (1 file), lme4 (1 file), lmerTest (1 file), Matplotlib (1 file), NiBabel (1 file), NumPy (1 file), pandas (1 file), patchwork (1 file), psych (1 file), rstatix (1 file), SciPy (1 file), tidyverse (1 file)
Availability: 1 check, the latest on 27 September 2026: the link answers
  • 27 September 2026: the link answers
6 files, not copied: shown from their source

OSCR keeps no copy of these files: this repository has no license that allows it. The reader above shows each one from its source, fetched by your browser at commit b1f8280, when its fingerprint is the one OSCR verified. How this works.

The paper's code and data availability statement is in the Data section.

Tracing map

Proposed by the machine: these links were found in the paper and verified at the source, without human review. The map will receive a Zenodo DOI once one of the paper's authors has validated it with their ORCID.

What the map holds:

  • 4 repositories of the authors' code, each at its verified commit, with its license and how the link was found in the paper;
  • 28 scripts, each with its path and the digest of its content;
  • 9 matches between paragraphs of the paper and lines of the code (method lexical-v1);
  • neither the text of the paper nor the code itself.

Its JSON (tracing-map.json) is deposited on Zenodo with its DOI once the map is validated.

Data

Datasets cited

Code and data availability statement

The paper has a code and data availability statement. Its license (CC BY-NC-ND) does not allow reproducing it here; in short, from what the harvester recognized in it:

  • no repository, dataset or request procedure was recognized in it

Read it in the paper: doi.org/10.1073/pnas.2603114123.

Versions

The history of this record: each version stored by the harvester or made by a correction of its authors or of the maintainers of its code, and what changed in its facts. The texts of the paper (its abstract, its availability statements) are not part of it; versions that changed only those are not listed.

Version 1, 27 September 2026: the first record

Recorded: type, language, journal, volume, issue, pages, dates, 4 authors, 5 keywords, 12 MeSH terms, 5 funders, 123 references.

Cite

This paper

Ilyés, A., Brosig, B. M., Mező, G., & Keresztes, A. (2026). The human hippocampus can pattern separate memories by meaning. Proceedings of the National Academy of Sciences of the United States of America, 123(31), e2603114123. https://doi.org/10.1073/pnas.2603114123

BibTeX

@article{ilyes2026human,
author = {Ilyés, Alex and Brosig, Borbála Mónika and Mező, György and Keresztes, Attila},
title = {{The human hippocampus can pattern separate memories by meaning}},
journal = {Proceedings of the National Academy of Sciences of the United States of America},
year = {2026},
month = jul,
volume = {123},
number = {31},
pages = {e2603114123},
publisher = {National Academy of Sciences},
issn = {0027-8424},
doi = {10.1073/pnas.2603114123},
url = {https://doi.org/10.1073/pnas.2603114123},
pmid = {42520121},
pmcid = {PMC13438472}
}

RIS

TY - JOUR
AU - Ilyés, Alex
AU - Brosig, Borbála Mónika
AU - Mező, György
AU - Keresztes, Attila
TI - The human hippocampus can pattern separate memories by meaning
T2 - Proceedings of the National Academy of Sciences of the United States of America
J2 - Proc Natl Acad Sci U S A
PY - 2026
DA - 2026/07/28
VL - 123
IS - 31
SP - e2603114123
SN - 0027-8424
PB - National Academy of Sciences
DO - 10.1073/pnas.2603114123
UR - https://doi.org/10.1073/pnas.2603114123
LA - en
ER -

CSL-JSON

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