Diffusion-relaxation MRI as virtual histology: separable microstructural signatures of AD pathology in ex vivo human brain

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• The repository at the source (pyushkevich/greedy)
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1. #!/bin/bash
2. set -x -e
4. pip install --upgrade pip setuptools wheel cmake ninja
6. echo "Check CMAKE version"
7. cmake --version
9. mkdir -p be/install && cd be
11. echo "Checking folder structure"
12. ls -lh .
13. ls -lh ..
15. # Download and build VTK
16. LIB_LOCATION=build
17. if [[ $1 =~ ubuntu-.* ]]; then
18. VTK_BINARY=vtk-wheel-sdk-9.3.1-cp310-cp310-manylinux_2_17_x86_64.manylinux2014_x86_64.tar.xz
19. DYLD_SUFFIX=so
20. MAKEFLAGS="-- -j 8"
21. elif [[ $1 == macos-13 ]]; then
22. VTK_BINARY=vtk-wheel-sdk-9.3.1-cp310-cp310-macosx_10_10_x86_64.tar.xz
23. DYLD_SUFFIX=dylib
24. MAKEFLAGS="-- -j 8"
25. elif [[ $1 == macos-14 ]]; then
26. VTK_BINARY=vtk-wheel-sdk-9.3.1-cp310-cp310-macosx_11_0_arm64.tar.xz
27. DYLD_SUFFIX=dylib
28. elif [[ $1 =~ windows-.* ]]; then
29. VTK_BINARY=vtk-wheel-sdk-9.3.1-cp310-cp310-win_amd64.tar.xz
30. DYLD_SUFFIX=dll
31. LIB_LOCATION=bin
32. CMAKE_RELEASE_COMMAND="--config Release"
33. else
34. exit 255
35. fi
37. # Install Eigen
38. git clone -b 3.4.0 https://gitlab.com/libeigen/eigen.git
39. cmake \
40. -DBUILD_EXAMPLES=OFF \
41. -DBUILD_TESTING=OFF \
42. -DCMAKE_BUILD_TYPE=Release \
43. -DCMAKE_INSTALL_PREFIX=./install \
44. -B eigen/build \
45. eigen
47. cmake --build eigen/build --target install $MAKEFLAGS $CMAKE_RELEASE_COMMAND
49. # Install VTK from binary wheels provided by Kitware
50. mkdir -p install/vtk install/vtk/shared
51. curl -L https://www.vtk.org/files/release/9.3/${VTK_BINARY} -o ./install/vtk/vtk-wheel-sdk.tar.xz
52. tar -xJvf ./install/vtk/vtk-wheel-sdk.tar.xz --strip-components 1 -C $PWD/install/vtk
53. ln $(find $PWD/install/vtk/${LIB_LOCATION} -name "*.${DYLD_SUFFIX}") install/vtk/shared
55. # Link the shared libraries needed for delocate into a simple directory
57. # Build ITK
58. git clone -b v5.2.1 https://github.com/InsightSoftwareConsortium/ITK.git ITK
59. cmake \
60. -DModule_MorphologicalContourInterpolation=ON \
61. -DBUILD_EXAMPLES=OFF \
62. -DBUILD_TESTING=OFF \
63. -DCMAKE_BUILD_TYPE=Release \
64. -DCMAKE_INSTALL_PREFIX=./install \
65. -DCMAKE_POSITION_INDEPENDENT_CODE=ON \
66. -DCMAKE_POLICY_VERSION_MINIMUM=3.5 \
67. -B ITK/build \
68. ITK
70. cmake --build ITK/build --target install $MAKEFLAGS $CMAKE_RELEASE_COMMAND
72. #git clone -b v9.3.1 https://github.com/Kitware/VTK.git VTK
73. #cmake \
74. # -DBUILD_EXAMPLES=OFF \
75. # -DBUILD_TESTING=OFF \
76. # -DCMAKE_BUILD_TYPE=Release \
77. # -DBUILD_SHARED_LIBS=OFF \
78. # -DVTK_REQUIRED_OBJCXX_FLAGS="" \
79. # -DCMAKE_INSTALL_PREFIX=./install \
80. # -B VTK/build \
81. # VTK
82. #cmake --build VTK/build --target install --config Release
84. # Build Greedy
85. # git clone -b master https://github.com/pyushkevich/greedy.git greedy
86. cmake \
87. -DCMAKE_BUILD_TYPE=Release \
88. -DCMAKE_INSTALL_PREFIX=./install \
89. -DGREEDY_BUILD_LMSHOOT=ON \
90. -DGREEDY_BUILD_WRAPPING=ON \
91. -DCMAKE_PREFIX_PATH="$PWD/install" \
92. -DCMAKE_POSITION_INDEPENDENT_CODE=ON \
93. -DVTK_DIR=$PWD/install/vtk/vtk-9.3.1.data/headers/cmake \
94. -B greedy/build \
95. ..
97. cmake --build greedy/build --target install $MAKEFLAGS $CMAKE_RELEASE_COMMAND

prebuild.sh at commit f5212c5, under Apache-2.0 · at the source

Overview

Authors: Eppu Manninen¹, Courtney J Comrie², Geidy E Serrano³, Thomas G Beach³, Elizabeth B Hutchinson², Dan Benjamini¹

1. National Institute on Aging, NIH
2. University of Arizona
3. Banner Sun Health Research Institute

Journal: Research square, pages rs.3.rs-9657671

Dates: published online 25 May 2026

Type: Research article · Language: English

License: CC BY

Identifiers: DOI · PMCID PMC13232456

Status: code verified

Categories: structural MRI / diffusion (modality), histology / microscopy (modality), human (organism), Alzheimer's / dementia (population), cellular / molecular (subfield)

Methods: Statistics, Machine learning, Preprocessing, Connectivity, fMRI & imaging

Keywords: Alzheimer’s disease, multidimensional MRI, myelin, microglia, phosphorylated tau, amyloid beta

Citations: 91 references in the paper

Abstract

Cognitive decline in Alzheimer's disease (AD) reflects progressive disruption of cellular and microstructural organization, yet the biological specificity of conventional MRI signals remains poorly understood. Multidimensional diffusion–relaxation MRI (MD-MRI) resolves sub-voxel tissue heterogeneity and may offer a framework for linking imaging signals to underlying neuropathology. We tested the hypothesis that neuronal, glial, and white matter pathologies in AD occupy separable regions of diffusion–relaxation space and generate spatially organized signatures associated with cognitive impairment. We integrated ex vivo MD-MRI with co-registered histology from 12 human donors spanning a range of Braak stages and pathological severity. Nested cross-validated elastic net models predicted voxelwise Aβ, pTau, microglia, and myelin burden from the multidimensional diffusion–relaxation density distribution. Regional associations were assessed across hippocampal subfields and white matter, and MRI-predicted pathology was related to ante-mortem Mini-Mental State Examination scores. Distinct diffusion–relaxation components were preferentially associated with different pathological markers, indicating separable microstructural signatures. MRI-derived predictions corresponded significantly with histological measures of myelin (ρ = 0.77), pTau (ρ = 0.62), and microglia (ρ = 0.61), with weaker correspondence for Aβ (ρ = 0.45). Regionally, predicted pathology recapitulated known patterns of selective vulnerability, with elevated pTau and microglial signal in hippocampal subfields and dominant myelin-associated signal in white matter (p < 0.0001). Higher predicted hippocampal pTau was strongly associated with worse cognitive performance (ρ = −0.88, p = 0.0014), with a moderate association in white matter (ρ = −0.66, p = 0.036). These findings demonstrate that AD-related pathological processes manifest as distinct, spatially organized diffusion–relaxation signatures, providing mechanistic insight into the microstructural basis of MRI contrasts. As clinically feasible MD-MRI protocols continue to emerge, translation of these signatures to in vivo imaging may enable more biologically informed assessment of neurodegeneration.

Reproduced under the paper's license (CC BY), from the paper cited above.

Repositories

Its files are read in the Code ↔ Paper reader above.

pyushkevich/greedy

License: Apache-2.0

State: the link answers, verified on 28 September 2026

Evidence: files inventoried

Commit: f5212c50d72f4ce511ec1a6e142c58b169de714e, 3 August 2026

Languages: MATLAB (83), C/C++ (52), Shell (5), C (3), Python (1)

Size: 309 files, 144 scripts

Software Heritage: archived

Found in: the text, “Registration of histological images to MRI”

Holds: README, license file, environment (pyproject.toml), continuous integration, documentation

Not found: CITATION.cff, tests

Tools: Image Processing Toolbox (12 files), Statistics and Machine Learning Toolbox (1 file), Tools for NIfTI and ANALYZE image (MATLAB) (1 file)

Availability: 1 check, the latest on 28 September 2026: the link answers

• 28 September 2026: the link answers

146 files

• .github/workflows/prebuild.sh, Shell, 97 lines
• docs/conf.py, Python, 172 lines
• matlab/diff/depthmap/depthmap.m, MATLAB, 15 lines
• matlab/diff/depthmap/runme.m, MATLAB, 4 lines
• matlab/diff/diff_objective.m, MATLAB, 61 lines
• matlab/diff/diff_objective_gateux_deriv.m, MATLAB, 48 lines
• matlab/diff/diff_objective_simple.m, MATLAB, 42 lines
• matlab/diff/diffopmat.m, MATLAB, 27 lines
• matlab/diff/diffopt.m, MATLAB, 57 lines
• matlab/diff/diffplot.m, MATLAB, 51 lines
• matlab/diff/elcallback.m, MATLAB, 89 lines
• matlab/diff/elevolve.m, MATLAB, 79 lines
• matlab/diff/elupdate.m, MATLAB, 26 lines
• matlab/diff/expmap.m, MATLAB, 24 lines
• matlab/diff/expmap_euler.m, MATLAB, 16 lines
• matlab/diff/expmap_gateaux_deriv.m, MATLAB, 53 lines
• matlab/diff/expmap_semi_lagrange.m, MATLAB, 27 lines
• matlab/diff/external/minFunc/ArmijoBacktrack.m, MATLAB, 134 lines
• matlab/diff/external/minFunc/WolfeLineSearch.m, MATLAB, 378 lines
• matlab/diff/external/minFunc/autoGrad.m, MATLAB, 1 line
• matlab/diff/external/minFunc/autoHess.m, MATLAB, 1 line
• matlab/diff/external/minFunc/autoHv.m, MATLAB, 11 lines
• matlab/diff/external/minFunc/autoTensor.m, MATLAB, 1 line
• matlab/diff/external/minFunc/conjGrad.m, MATLAB, 70 lines
• matlab/diff/external/minFunc/example_minFunc.m, MATLAB, 59 lines
• matlab/diff/external/minFunc/lbfgs.m, MATLAB, 40 lines
• matlab/diff/external/minFunc/lbfgsC.c, C, 115 lines
• matlab/diff/external/minFunc/lbfgsUpdate.m, MATLAB, 20 lines
• matlab/diff/external/minFunc/mchol.m, MATLAB, 60 lines
• matlab/diff/external/minFunc/mcholC.c, C, 199 lines
• matlab/diff/external/minFunc/mcholinc.m, MATLAB, 25 lines
• matlab/diff/external/minFunc/minFunc.m, MATLAB, 769 lines
• matlab/diff/external/minFunc/minFuncBC.m, MATLAB, 285 lines
• matlab/diff/external/minFunc/minFuncEC.m, MATLAB, 66 lines
• matlab/diff/external/minFunc/minFuncIC.m, MATLAB, 76 lines
• matlab/diff/external/minFunc/minFunc_processInputOptions.m, MATLAB, 121 lines
• matlab/diff/external/minFunc/polyinterp.m, MATLAB, 140 lines
• matlab/diff/external/minFunc/precondDiag.m, MATLAB, 2 lines
• matlab/diff/external/minFunc/precondTriu.m, MATLAB, 2 lines
• matlab/diff/external/minFunc/precondTriuDiag.m, MATLAB, 2 lines
• matlab/diff/external/minFunc/taylorModel.m, MATLAB, 38 lines
• matlab/diff/generate_test_fft.m, MATLAB, 77 lines
• matlab/diff/gridplot.m, MATLAB, 21 lines
• matlab/diff/image_downsample.m, MATLAB, 10 lines
• matlab/diff/image_upsample.m, MATLAB, 10 lines
• matlab/diff/lddmm_basic.m, MATLAB, 37 lines
• matlab/diff/lddmm_compose_diffeomorphisms.m, MATLAB, 13 lines
• matlab/diff/lddmm_diff_variation.m, MATLAB, 1 line
• matlab/diff/lddmm_example_cee_to_circ.m, MATLAB, 13 lines
• matlab/diff/lddmm_hilbert_dot_product.m, MATLAB, 9 lines
• matlab/diff/lddmm_init.m, MATLAB, 50 lines
• matlab/diff/lddmm_integrate_field_and_jacobian_semi_lagrangian.m, MATLAB, 105 lines
• matlab/diff/lddmm_integrate_field_euler.m, MATLAB, 22 lines
• matlab/diff/lddmm_integrate_field_semi_lagrangian.m, MATLAB, 45 lines
• matlab/diff/lddmm_jacobian_determinant.m, MATLAB, 10 lines
• matlab/diff/lddmm_objective_and_gateaux.m, MATLAB, 65 lines
• matlab/diff/lddmm_objective_and_gradient.m, MATLAB, 65 lines
• matlab/diff/lddmm_optimize.m, MATLAB, 41 lines
• matlab/diff/lddmm_optimize_basic.m, MATLAB, 52 lines
• matlab/diff/lddmm_optimize_lbgfs.m, MATLAB, 30 lines
• matlab/diff/lddmm_phi_gateaux.m, MATLAB, 3 lines
• matlab/diff/lddmm_phi_s_t.m, MATLAB, 1 line
• matlab/diff/lddmm_plot_field.m, MATLAB, 9 lines
• matlab/diff/lddmm_plot_greens_function.m, MATLAB, 14 lines
• matlab/diff/lddmm_plot_problem.m, MATLAB, 58 lines
• matlab/diff/lddmm_plot_problem_simple.m, MATLAB, 37 lines
• matlab/diff/lddmm_random_field.m, MATLAB, 55 lines
• matlab/diff/lddmm_register_multiscale.m, MATLAB, 37 lines
• matlab/diff/lddmm_reparameterize_vector_field.m, MATLAB, 46 lines
• matlab/diff/lddmm_resample_in_time.m, MATLAB, 11 lines
• matlab/diff/lddmm_test_gradient.m, MATLAB, 52 lines
• matlab/diff/lddmm_test_inverse_consistency.m, MATLAB, 37 lines
• matlab/diff/lddmm_trivial_objective_and_gradient.m, MATLAB, 51 lines
• matlab/diff/lddmm_vector_field_dot_product.m, MATLAB, 17 lines
• matlab/diff/lddmm_warp_scalar_field.m, MATLAB, 10 lines
• matlab/diff/mkgridimage.m, MATLAB, 5 lines
• matlab/diff/normed_dot_product.m, MATLAB, 6 lines
• matlab/diff/test_lddmm.m, MATLAB, 21 lines
• matlab/diff/test_lddmm_brains.m, MATLAB, 24 lines
• matlab/diff/test_lddmm_corpus.m, MATLAB, 21 lines
• matlab/diff/testgateaux.m, MATLAB, 45 lines
• matlab/diff/testinvmap.m, MATLAB, 26 lines
• matlab/diff/warp.m, MATLAB, 4 lines
• matlab/ncc_test/compute_ncc.m, MATLAB, 20 lines
• matlab/ncc_test/compute_wncc.m, MATLAB, 23 lines
• matlab/ncc_test/ncc_test_feb2021.m, MATLAB, 68 lines
• matlab/ncc_test/weighted_ncc_test_feb2021.m, MATLAB, 110 lines
• scripts/template/greedy_build_template.sh, Shell, 591 lines
• scripts/template/pybatch.sh, Shell, 121 lines
• src/AffineCostFunctions.h, C/C++, 457 lines
• src/AffineTransformUtilities.h, C/C++, 231 lines
• src/CommandLineHelper.h, C/C++, 420 lines
• src/DifferentiableScalingAndSquaring.h, C/C++, 194 lines
• src/GreedyAPI.h, C/C++, 368 lines
• src/GreedyException.h, C/C++, 70 lines
• src/GreedyMeshIO.h, C/C++, 28 lines
• src/GreedyParameters.h, C/C++, 507 lines
• src/ITKFilters/include/CompositeImageNanMaskingFilter.h, C/C++, 104 lines
• src/ITKFilters/include/FastLinearInterpolator.h, C/C++, 1,136 lines
• src/ITKFilters/include/FastWarpCompositeImageFilter.h, C/C++, 160 lines
• src/ITKFilters/include/ImageRegionConstIteratorWithIndexOverride.h, C/C++, 78 lines
• src/ITKFilters/include/JacobianDeterminantImageFilter.h, C/C++, 86 lines
• src/ITKFilters/include/LieBracketFilter.h, C/C++, 99 lines
• src/ITKFilters/include/LinearTransformToWarpFilter.h, C/C++, 101 lines
• src/ITKFilters/include/MahalanobisDistanceToTargetWarpMetric.h, C/C++, 134 lines
• src/ITKFilters/include/MultiComponentApproximateNCCImageMetric.h, C/C++, 235 lines
• src/ITKFilters/include/MultiComponentImageMetricBase.h, C/C++, 326 lines
• src/ITKFilters/include/MultiComponentMetricReport.h, C/C++, 70 lines
• src/ITKFilters/include/MultiComponentMutualInfoImageMetric.h, C/C++, 206 lines
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• src/ITKFilters/include/MultiComponentQuantileBasedNormalizationFilter.h, C/C++, 198 lines
• src/ITKFilters/include/MultiComponentWeightedNCCImageMetric.h, C/C++, 181 lines
• src/ITKFilters/include/MultiImageOpticalFlowImageFilter.h, C/C++, 162 lines
• src/ITKFilters/include/OneDimensionalInPlaceAccumulateFilter.h, C/C++, 119 lines
• src/ITKFilters/include/itkSLICSuperVoxelImageFilter.h, C/C++, 114 lines
• src/ITKFilters/include/itkVectorImageCentralDifferenceImageFunction.h, C/C++, 150 lines
• src/MultiImageRegistrationHelper.h, C/C++, 368 lines
• src/SparseSolvers/MKLSolverInterface.h, C/C++, 72 lines
• src/SparseSolvers/PardisoInterface.h, C/C++, 114 lines
• src/SparseSolvers/SparseMatrix.h, C/C++, 272 lines
• src/SparseSolvers/SparseSolver.h, C/C++, 45 lines
• src/SparseSolvers/SparseSolverException.h, C/C++, 20 lines
• src/SparseSolvers/TaucsInterface.h, C/C++, 68 lines
• src/TetraMeshConstraints.h, C/C++, 126 lines
• src/WarpFunctors.h, C/C++, 222 lines
• src/dijkstra/BinaryHeap.h, C/C++, 274 lines
• src/dijkstra/ShortestPath.h, C/C++, 266 lines
• src/lddmm_common.h, C/C++, 70 lines
• src/lddmm_data.h, C/C++, 467 lines
• src/lmshoot/PointSetGeodesicShooting.h, C/C++, 90 lines
• src/lmshoot/PointSetGeodesicToWarp.h, C/C++, 66 lines
• src/lmshoot/PointSetHamiltonianSystem.h, C/C++, 215 lines
• src/lmshoot/PointSetOptimalControlSystem.h, C/C++, 127 lines
• src/lmshoot/VTKArrays.h, C/C++, 61 lines
• src/lmshoot/ctpl_stl.h, C/C++, 251 lines
• src/multichunk/MultiChunkGreedy.h, C/C++, 26 lines
• src/propagation/PropagationAPI.h, C/C++, 121 lines
• src/propagation/PropagationIO.h, C/C++, 107 lines
• src/propagation/PropagationInputBuilder.h, C/C++, 115 lines
• src/propagation/PropagationTools.h, C/C++, 208 lines
• testing/data/runpair.sh, Shell, 32 lines
• testing/data/src/phantoms.sh, Shell, 35 lines
• utilities/hexdump.c, C, 2,107 lines
• utilities/hexdump.h, C/C++, 215 lines
• LICENSE, License, 202 lines
• README.md, Text, 5 lines

At the source: github.com/pyushkevich/greedy

dan-benjamini/madco

License: none: the authors keep all their rights

State: the link answers, verified on 28 September 2026

Evidence: files inventoried

Commit: d665140a31135349ec280e0a4adfd2f9b86f70f6, 21 May 2025

Languages: MATLAB (618), C (70), JavaScript (13), C/C++ (6)

Size: 1,207 files, 707 scripts

Software Heritage: not archived

Found in: “Data availability”

Holds: README, documentation

Not found: license file, CITATION.cff, environment file, tests, continuous integration

Tools: Parallel Computing Toolbox (1 file)

Availability: 1 check, the latest on 28 September 2026: the link answers

• 28 September 2026: the link answers

708 files

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• MatlabFunctions/cvx/structures/cvx_s_upper_hankel.m, MATLAB, 16 lines
• MatlabFunctions/madco2D/MultiDprocFun.m, MATLAB, 847 lines
• MatlabFunctions/madco2D/NESMA_filtering.m, MATLAB, 26 lines
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• MatlabFunctions/madco2D/find_method.m, MATLAB, 42 lines
• MatlabFunctions/madco2D/find_multi_method.m, MATLAB, 47 lines
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• MatlabFunctions/madco2D/geo_mean.m, MATLAB, 73 lines
• MatlabFunctions/madco2D/readMDmri.m, MATLAB, 154 lines
• MatlabFunctions/madco2D/read_2dseq_3D_DiffIsoAcqIRpath.m, MATLAB, 93 lines
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• MatlabFunctions/madco2D/read_2dseq_3D_TETIpath.m, MATLAB, 69 lines
• MatlabFunctions/madco2D/read_2dseq_3D_TEpath.m, MATLAB, 68 lines
• MatlabFunctions/madco2D/read_2dseq_3D_TIpath.m, MATLAB, 68 lines
• README.md, Text, 4 lines

At the source: github.com/dan-benjamini/madco/

The paper's code and data availability statement is in the Data section.

Tracing map

Proposed by the machine: these links were found in the paper and verified at the source, without human review. The map will receive a Zenodo DOI once one of the paper's authors has validated it with their ORCID.

What the map holds:

• 2 repositories of the authors' code, each at its verified commit, with its license and how the link was found in the paper;
• 851 scripts, each with its path and the digest of its content;
• no match between paragraphs and code yet;
• neither the text of the paper nor the code itself.

Its JSON (tracing-map.json) is deposited on Zenodo with its DOI once the map is validated.

Data

No dataset and no data link were found in the paper.

Data availability

The datasets generated and analyzed during the current study are available from the corresponding author upon request, subject to applicable data sharing agreements and conditions of reuse. MATLAB source code for preprocessing and MADCO data inversion is freely available at https://github.com/dan-benjamini/madco/.

Reproduced under the paper's license (CC BY), from the paper cited above.

Versions

The history of this record: each version stored by the harvester or made by a correction of its authors or of the maintainers of its code, and what changed in its facts. The texts of the paper (its abstract, its availability statements) are not part of it; versions that changed only those are not listed.

Version 1, 28 September 2026: the first record

Recorded: type, language, journal, pages, dates, 6 authors, 6 keywords, 90 references.

Cite

This paper

Manninen, E., Comrie, C. J., Serrano, G. E., Beach, T. G., Hutchinson, E. B., & Benjamini, D. (2026). Diffusion-relaxation MRI as virtual histology: separable microstructural signatures of AD pathology in ex vivo human brain. Research square, rs.3.rs-9657671.

BibTeX

@article{manninen2026diffusion,
author = {Manninen, Eppu and Comrie, Courtney J and Serrano, Geidy E and Beach, Thomas G and Hutchinson, Elizabeth B and Benjamini, Dan},
title = {{Diffusion-relaxation MRI as virtual histology: separable microstructural signatures of AD pathology in ex vivo human brain}},
journal = {Research square},
year = {2026},
month = may,
pages = {rs.3.rs--9657671},
publisher = {American Journal Experts},
pmcid = {PMC13232456}
}

RIS

TY - JOUR
AU - Manninen, Eppu
AU - Comrie, Courtney J
AU - Serrano, Geidy E
AU - Beach, Thomas G
AU - Hutchinson, Elizabeth B
AU - Benjamini, Dan
TI - Diffusion-relaxation MRI as virtual histology: separable microstructural signatures of AD pathology in ex vivo human brain
T2 - Research square
PY - 2026
DA - 2026/05/25
SP - rs.3.rs
EP - 9657671
PB - American Journal Experts
LA - en
ER -

CSL-JSON

{
"id": "pmcid:PMC13232456",
"type": "article-journal",
"title": "Diffusion-relaxation MRI as virtual histology: separable microstructural signatures of AD pathology in ex vivo human brain",
"container-title": "Research square",
"author": [
{
"family": "Manninen",
"given": "Eppu"
},
{
"family": "Comrie",
"given": "Courtney J"
},
{
"family": "Serrano",
"given": "Geidy E"
},
{
"family": "Beach",
"given": "Thomas G"
},
{
"family": "Hutchinson",
"given": "Elizabeth B"
},
{
"family": "Benjamini",
"given": "Dan"
}
],
"page": "rs.3.rs-9657671",
"PMCID": "PMC13232456",
"publisher": "American Journal Experts",
"language": "en",
"issued": {
"date-parts": [
[
2026,
5,
25
]
]
}
}

The tracing map gets a citation of its own once an author has validated it and it has a DOI.

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The authors' code

• Code ↔ Paper, above
• matches not computed yet
• pyushkevich/greedy
• dan-benjamini/madco

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Licenses

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• the code: Apache-2.0
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