Neural substrates of ambiguity and beauty in haiku poetry: an fMRI study.
The 4 matches
- [1] § Materials and methods › fMRI data analysis › Psychophysiological interaction (PPI) analysis ↔ Matlab_script/run_ppi_firstlevel_batch_manualPPI.m, lines 102–143 · score 0.57 · PPI regressor, Tikhonov, GLM, signals, VOI, HRF
- [2] § Results › Brain results ↔ Matlab_script/mvpa_PCC_ambiguity.m, lines 157–205 · score 0.55 · linear SVM, cross validation, components, MVPA, PCC, ambiguity
- [3] § Results › Brain results ↔ Matlab_script/mvpa_LIPL_ambiguity.m, lines 157–204 · score 0.54 · linear SVM, cross validation, components, MVPA, ambiguity
- [4] § Materials and methods › fMRI data analysis › Multivoxel pattern analysis (MVPA) ↔ Matlab_script/mvpa_LIPL_ambiguity.m, lines 1–6 · score 0.50 · cross validation, left IPL, MVPA, SVMs, classified, ambiguity
Paper
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The authors' code
MATLAB · 414 lines · 16 KB · no license · 2 matches
- %% MVPA Step 3: LIPL ROI × Ambiguity Classification
- % 目的: Left IPLが曖昧度(low vs high)を分類できるかを検証
- % 方法: Leave-one-trial-out cross-validation + PCA + 線形SVM
- % Statistics Toolbox不要
- clear; clc;
- %% パラメータ設定
- TR = 2; % 秒
- peak_trs_offset = [2, 3, 4]; % onset後の秒数(4-10秒)
- roi_radius = 6; % mm
- roi_coord_mni = [-60, -42, 40]; % Left IPL
- n_pca_components = 10; % PCA次元数
- % パス設定
- base_dir = 'D:/mri_toolbox';
- csv_path = fullfile(base_dir, 'mri_time', 'beauty_time.csv');
- output_dir = fullfile(base_dir, 'MVPA_Results', 'LIPL_Ambiguity');
- if ~exist(output_dir, 'dir'), mkdir(output_dir); end
- % 被験者リスト
- all_ids = setdiff(1:44, [1, 2, 7, 28, 33]); % 39名
- %% CSVデータ読み込み
- fprintf('CSVデータ読み込み中...\n');
- opts = detectImportOptions(csv_path, 'Encoding', 'UTF-8');
- behavior_data = readtable(csv_path, opts);
- fprintf('✓ CSV読み込み完了\n\n');
- %% セッション番号マッピング
- session_map = containers.Map('KeyType', 'double', 'ValueType', 'double');
- session_map(9) = 6; session_map(43) = 7; session_map(6) = 6;
- session_map(37) = 6; session_map(18) = 7;
- %% 全被験者でループ
- results = struct();
- success_count = 0;
- fail_count = 0;
- for idx = 1:length(all_ids)
- sub_id = all_ids(idx);
- fprintf('========================================\n');
- fprintf('Processing Sub%02d (%d/%d)\n', sub_id, idx, length(all_ids));
- fprintf('========================================\n');
- try
- %% 1. 機能画像のパス取得
- if isKey(session_map, sub_id)
- session = session_map(sub_id);
- else
- session = 5;
- end
- func_dir = sprintf('%s/HAIKU_%02d/%d_mb_ep_bold_mb4_sl76_tr2_task2/', ...
- base_dir, sub_id, session);
- func_files = spm_select('FPList', func_dir, '^s.*\.nii$');
- if isempty(func_files)
- error('機能画像が見つかりません');
- end
- fprintf('✓ 機能画像: %d ボリューム\n', size(func_files, 1));
- %% 2. 被験者の行動データ抽出
- sub_data = behavior_data(strcmp(strtrim(string(behavior_data.ID)), num2str(sub_id)) | ...
- strcmp(strtrim(string(behavior_data.ID)), sprintf('%02d', sub_id)), :);
- if isempty(sub_data)
- error('CSVにデータが見つかりません');
- end
- n_trials = height(sub_data);
- fprintf('✓ 試行数: %d\n', n_trials);
- % Ambiguityラベル(1=low, 2=high)
- ambiguity_labels = sub_data.condition;
- % 各条件の試行数を確認
- n_low = sum(ambiguity_labels == 1);
- n_high = sum(ambiguity_labels == 2);
- fprintf(' Low ambiguity: %d trials\n', n_low);
- fprintf(' High ambiguity: %d trials\n', n_high);
- if n_low < 2 || n_high < 2
- error('各条件に最低2試行必要です');
- end
- %% 3. ROIマスク作成
- V = spm_vol(deblank(func_files(1,:)));
- % MNI → voxel座標変換
- vox_coord = round(inv(V.mat) * [roi_coord_mni'; 1]);
- vox_coord = vox_coord(1:3);
- fprintf('✓ ROI中心 (MNI): [%d, %d, %d]\n', roi_coord_mni);
- fprintf('✓ ROI中心 (vox): [%d, %d, %d]\n', vox_coord);
- % 6mm球内のボクセルを特定
- [X, Y, Z] = ndgrid(1:V.dim(1), 1:V.dim(2), 1:V.dim(3));
- voxel_coords = [X(:), Y(:), Z(:)];
- % 各ボクセルのMNI座標を計算
- mni_coords = V.mat * [voxel_coords'; ones(1, size(voxel_coords, 1))];
- mni_coords = mni_coords(1:3, :)';
- % ROI中心からの距離
- distances = sqrt(sum((mni_coords - repmat(roi_coord_mni, size(mni_coords, 1), 1)).^2, 2));
- roi_mask = distances <= roi_radius;
- n_voxels = sum(roi_mask);
- fprintf('✓ ROI内ボクセル数: %d\n', n_voxels);
- %% 4. 各試行のBOLD信号を抽出(複数TRの平均)
- X_data = zeros(n_trials, n_voxels); % 試行 × ボクセル
- for trial = 1:n_trials
- onset_sec = sub_data.time_b(trial);
- % 複数TRを平均
- trial_signal = zeros(length(peak_trs_offset), n_voxels);
- for t_idx = 1:length(peak_trs_offset)
- peak_sec = onset_sec + peak_trs_offset(t_idx);
- peak_tr = round(peak_sec / TR) + 1;
- if peak_tr > size(func_files, 1)
- peak_tr = size(func_files, 1);
- end
- vol = spm_read_vols(spm_vol(deblank(func_files(peak_tr,:))));
- roi_voxels = vol(roi_mask);
- trial_signal(t_idx, :) = roi_voxels(:)';
- end
- % 複数TRの平均
- X_data(trial, :) = mean(trial_signal, 1);
- end
- fprintf('✓ BOLD信号抽出完了: [%d trials × %d voxels]\n', n_trials, n_voxels);
- %% 5. PCAで次元削減
- n_components = min(n_pca_components, n_trials - 2);
- X_centered = X_data - mean(X_data, 1);
- % SVDでPCA
- [U, S, V_pca] = svd(X_centered', 'econ');
- pca_components = U(:, 1:n_components);
- % 説明分散
- explained_var = diag(S).^2 / sum(diag(S).^2);
- cumsum_var = cumsum(explained_var(1:n_components));
- fprintf('✓ PCAで%dボクセル → %d次元に削減\n', n_voxels, n_components);
- fprintf(' 累積説明分散: %.1f%%\n', cumsum_var(end) * 100);
- %% 6. Leave-One-Trial-Out Cross-Validation(線形SVM)
- predictions = zeros(n_trials, 1);
- decision_values = zeros(n_trials, 1);
- for trial = 1:n_trials
- % 訓練データとテストデータを分割
- train_idx = setdiff(1:n_trials, trial);
- test_idx = trial;
- X_train = X_data(train_idx, :);
- y_train = ambiguity_labels(train_idx);
- X_test = X_data(test_idx, :);
- % 訓練データで中心化
- train_mean = mean(X_train, 1);
- % PCA変換
- X_train_centered = X_train - train_mean;
- X_test_centered = X_test - train_mean;
- X_train_pca = X_train_centered * pca_components;
- X_test_pca = X_test_centered * pca_components;
- % 線形SVM(手動実装)
- % 2クラス分類: y = 1 or 2 → -1 or +1 に変換
- y_train_svm = y_train;
- y_train_svm(y_train == 1) = -1; % Low ambiguity
- y_train_svm(y_train == 2) = +1; % High ambiguity
- % Linear SVM (simplified version using ridge regression)
- lambda = 1.0;
- XtX = X_train_pca' * X_train_pca;
- XtX_reg = XtX + lambda * eye(size(XtX));
- w = XtX_reg \ (X_train_pca' * y_train_svm);
- % 予測(決定値)
- decision_val = X_test_pca * w;
- decision_values(trial) = decision_val;
- % 分類(符号で判定)
- if decision_val >= 0
- predictions(trial) = 2; % High ambiguity
- else
- predictions(trial) = 1; % Low ambiguity
- end
- end
- fprintf('✓ Cross-validation完了\n');
- %% 7. 精度計算
- y_true = ambiguity_labels;
- y_pred = predictions;
- % 正解率(Accuracy)
- accuracy = sum(y_true == y_pred) / n_trials;
- % 各条件の正解率
- low_idx = (y_true == 1);
- high_idx = (y_true == 2);
- accuracy_low = sum(y_true(low_idx) == y_pred(low_idx)) / sum(low_idx);
- accuracy_high = sum(y_true(high_idx) == y_pred(high_idx)) / sum(high_idx);
- % Balanced accuracy
- balanced_accuracy = (accuracy_low + accuracy_high) / 2;
- % Confusion matrix
- tp = sum(y_true == 2 & y_pred == 2);
- tn = sum(y_true == 1 & y_pred == 1);
- fp = sum(y_true == 1 & y_pred == 2);
- fn = sum(y_true == 2 & y_pred == 1);
- % Sensitivity & Specificity
- if (tp + fn) > 0
- sensitivity = tp / (tp + fn);
- else
- sensitivity = NaN;
- end
- if (tn + fp) > 0
- specificity = tn / (tn + fp);
- else
- specificity = NaN;
- end
- % Binomial test p-value
- n_correct = sum(y_true == y_pred);
- p_binomial = binomial_test(n_correct, n_trials, 0.5);
- fprintf('\n--- デコーディング結果 ---\n');
- fprintf('Accuracy: %.2f%% (%d/%d)\n', accuracy * 100, sum(y_true == y_pred), n_trials);
- fprintf('Balanced Accuracy: %.2f%%\n', balanced_accuracy * 100);
- fprintf(' Low ambiguity正解率: %.2f%% (%d/%d)\n', accuracy_low * 100, sum(y_true(low_idx) == y_pred(low_idx)), sum(low_idx));
- fprintf(' High ambiguity正解率: %.2f%% (%d/%d)\n', accuracy_high * 100, sum(y_true(high_idx) == y_pred(high_idx)), sum(high_idx));
- fprintf('Sensitivity: %.2f%%\n', sensitivity * 100);
- fprintf('Specificity: %.2f%%\n', specificity * 100);
- fprintf('p = %.4f (binomial test vs chance)\n', p_binomial);
- fprintf('\nConfusion Matrix:\n');
- fprintf(' Predicted Low Predicted High\n');
- fprintf('Actual Low %3d %3d\n', tn, fp);
- fprintf('Actual High %3d %3d\n', fn, tp);
- %% 8. 結果を保存
- results(idx).sub_id = sub_id;
- results(idx).n_trials = n_trials;
- results(idx).n_low = n_low;
- results(idx).n_high = n_high;
- results(idx).n_voxels = n_voxels;
- results(idx).n_components = n_components;
- results(idx).explained_var = cumsum_var(end);
- results(idx).accuracy = accuracy;
- results(idx).balanced_accuracy = balanced_accuracy;
- results(idx).accuracy_low = accuracy_low;
- results(idx).accuracy_high = accuracy_high;
- results(idx).sensitivity = sensitivity;
- results(idx).specificity = specificity;
- results(idx).p_value = p_binomial;
- results(idx).confusion_matrix = [tn, fp; fn, tp];
- results(idx).predictions = predictions;
- results(idx).true_labels = y_true;
- results(idx).decision_values = decision_values;
- % 個別ファイルに保存
- sub_result = results(idx);
- save(fullfile(output_dir, sprintf('sub%02d_result.mat', sub_id)), 'sub_result');
- fprintf('✓ 保存完了\n\n');
- success_count = success_count + 1;
- catch ME
- fprintf('✗ エラー: %s\n', ME.message);
- if ~isempty(ME.stack)
- fprintf(' スタック: %s (line %d)\n\n', ME.stack(1).name, ME.stack(1).line);
- end
- fail_count = fail_count + 1;
- results(idx).sub_id = sub_id;
- results(idx).error = ME.message;
- end
- end
- %% 9. グループレベル集計
- fprintf('========================================\n');
- fprintf('全被験者処理完了\n');
- fprintf('========================================\n');
- fprintf('成功: %d / %d\n', success_count, length(all_ids));
- fprintf('失敗: %d / %d\n\n', fail_count, length(all_ids));
- % 成功した被験者の結果を集計
- valid_idx = arrayfun(@(x) isfield(x, 'accuracy'), results);
- valid_results = results(valid_idx);
- if isempty(valid_results)
- fprintf('有効な結果がありません\n');
- return;
- end
- all_accuracies = [valid_results.accuracy];
- all_balanced_accuracies = [valid_results.balanced_accuracy];
- all_p_values = [valid_results.p_value];
- fprintf('--- グループレベル統計 ---\n');
- fprintf('有効被験者数: %d\n', length(all_accuracies));
- fprintf('\n【Accuracy】\n');
- fprintf('平均: %.2f%% (SD = %.2f%%)\n', mean(all_accuracies) * 100, std(all_accuracies) * 100);
- fprintf('中央値: %.2f%%\n', median(all_accuracies) * 100);
- fprintf('範囲: [%.2f%%, %.2f%%]\n', min(all_accuracies) * 100, max(all_accuracies) * 100);
- fprintf('\n【Balanced Accuracy】\n');
- fprintf('平均: %.2f%% (SD = %.2f%%)\n', mean(all_balanced_accuracies) * 100, std(all_balanced_accuracies) * 100);
- fprintf('中央値: %.2f%%\n', median(all_balanced_accuracies) * 100);
- % 分布
- fprintf('\n【分布】\n');
- fprintf('Accuracy > 60%%: %d名 (%.1f%%)\n', sum(all_accuracies > 0.6), sum(all_accuracies > 0.6)/length(all_accuracies)*100);
- fprintf('Accuracy > 55%%: %d名 (%.1f%%)\n', sum(all_accuracies > 0.55), sum(all_accuracies > 0.55)/length(all_accuracies)*100);
- fprintf('Accuracy > 50%%: %d名 (%.1f%%)\n', sum(all_accuracies > 0.5), sum(all_accuracies > 0.5)/length(all_accuracies)*100);
- fprintf('Accuracy < 50%%: %d名 (%.1f%%)\n', sum(all_accuracies < 0.5), sum(all_accuracies < 0.5)/length(all_accuracies)*100);
- % 有意な被験者数
- sig_count = sum(all_p_values < 0.05);
- fprintf('\n有意な被験者数 (p<0.05): %d / %d (%.1f%%)\n', sig_count, length(all_p_values), sig_count/length(all_p_values)*100);
- % One-sample t-test
- mean_acc = mean(all_accuracies);
- std_acc = std(all_accuracies);
- n = length(all_accuracies);
- chance_level = 0.5;
- t_group = (mean_acc - chance_level) / (std_acc / sqrt(n));
- fprintf('\n【グループレベル検定】\n');
- fprintf('平均 Accuracy vs Chance (50%%):\n');
- fprintf('t(%d) = %.3f\n', n-1, t_group);
- if n > 30
- p_group = 1 - normcdf_manual(abs(t_group));
- fprintf('p ≈ %.6f (one-tailed)\n', p_group);
- if p_group < 0.001
- fprintf('*** p < 0.001\n');
- elseif p_group < 0.01
- fprintf('** p < 0.01\n');
- elseif p_group < 0.05
- fprintf('* p < 0.05\n');
- end
- end
- % 効果量
- cohens_d = (mean_acc - chance_level) / std_acc;
- fprintf('Cohen''s d = %.3f\n', cohens_d);
- % 95%信頼区間
- se_acc = std_acc / sqrt(n);
- ci_lower = mean_acc - 1.96 * se_acc;
- ci_upper = mean_acc + 1.96 * se_acc;
- fprintf('95%% CI: [%.2f%%, %.2f%%]\n', ci_lower * 100, ci_upper * 100);
- %% 10. 結果を保存
- save(fullfile(output_dir, 'group_results.mat'), 'results', 'all_accuracies', 'all_balanced_accuracies', 'all_p_values');
- % サマリーをテキストファイルにも保存
- fid = fopen(fullfile(output_dir, 'summary.txt'), 'w');
- fprintf(fid, 'MVPA Results: LIPL × Ambiguity Classification\n');
- fprintf(fid, '==============================================\n\n');
- fprintf(fid, 'パラメータ:\n');
- fprintf(fid, ' ROI: Left IPL [-60, -42, 40], 6mm球\n');
- fprintf(fid, ' 時間窓: onset + [%s]秒の平均\n', num2str(peak_trs_offset));
- fprintf(fid, ' PCA次元: %d\n\n', n_pca_components);
- fprintf(fid, '成功: %d / %d\n', success_count, length(all_ids));
- fprintf(fid, '失敗: %d / %d\n\n', fail_count, length(all_ids));
- fprintf(fid, '平均 Accuracy: %.2f%% (SD = %.2f%%)\n', mean_acc * 100, std_acc * 100);
- fprintf(fid, '中央値: %.2f%%\n', median(all_accuracies) * 100);
- fprintf(fid, '範囲: [%.2f%%, %.2f%%]\n\n', min(all_accuracies) * 100, max(all_accuracies) * 100);
- fprintf(fid, 't(%d) = %.3f\n', n-1, t_group);
- fprintf(fid, 'Cohen''s d = %.3f\n', cohens_d);
- fprintf(fid, '95%% CI: [%.2f%%, %.2f%%]\n', ci_lower * 100, ci_upper * 100);
- fclose(fid);
- fprintf('\n✓ 全結果を保存: %s\n', output_dir);
- %% ヘルパー関数
- function p = normcdf_manual(x)
- t = 1 ./ (1 + 0.2316419 * abs(x));
- d = 0.3989423 * exp(-x.*x/2);
- p = d .* t .* (0.3193815 + t .* (-0.3565638 + t .* (1.781478 + t .* (-1.821256 + t .* 1.330274))));
- p(x > 0) = 1 - p(x > 0);
- end
- function p = binomial_test(k, n, p0)
- % 正規近似版(簡易)
- expected = n * p0;
- variance = n * p0 * (1 - p0);
- z = (k - expected) / sqrt(variance);
- p = 2 * (1 - normcdf_manual(abs(z)));
- p = min(p, 1);
- end
mvpa_LIPL_ambiguity.m, no license · at the source
Overview
- Experimental Psychology Unit, Faculty of Humanities and Social Sciences, Helmut Schmidt University/University of the Federal Armed Forces Hamburg, Hamburg, Germany
- Japan Society for the Promotion of Science, Tokyo, Japan
- Graduate School of Education, Kyoto University, Kyoto, Japan
Abstract
Although neuroaesthetics has extensively examined visual art and music, the neural mechanisms underlying poetic appreciation remain less explored. This fMRI study investigated how the brain processes ambiguity and aesthetic experiences using haiku, the world’s shortest form of poetry. Originating in Japan, haiku is characterized by extreme brevity, often omitting information to compel readers to generate mental imagery and confront inherent ambiguities. Thirty-nine participants read haiku, which were pre-categorized via an independent survey into high- and low-ambiguity conditions, and rated their beauty inside the scanner. Consistent with previous findings, the behavioral results indicated a preference for low-ambiguity haiku (a small effect, standardized β = −0.16). Neuroimaging revealed that reading low-ambiguity haiku, compared to high-ambiguity ones, elicited greater activation in the posterior cingulate cortex and the left inferior parietal lobule, including the angular gyrus (cluster-level pFWE = 0.033 and 0.005, respectively). These regions are associated with the default mode network and semantic integration, suggesting that more interpretable poems may involve stronger self-referential processing and scene construction. Higher beauty ratings were significantly associated with increased activation in the primary visual cortex (cluster-level pFWE = 0.002). Our findings imply that the aesthetic appeal of haiku depends on semantic accessibility, with reduced ambiguity associated with greater engagement of self-referential processing in the default mode network and with increased activity in the visual cortex. More broadly, using haiku as a model offers a foundation for neuroscientific investigations of ambiguity and aesthetic experience across poetry, narratives, and other art forms.
Reproduced under the paper's license (CC BY), from the paper cited above.
Repository
Its files are read in the Code ↔ Paper reader above, with 4 matches between paragraphs and lines of code.
OSF emsp4
Availability: 1 check, the latest on 27 September 2026: the link answers (HTTP 200)
- 27 September 2026: the link answers (HTTP 200)
13 files
- Matlab_script/
mvpa_LIPL_ambiguity.m , MATLAB, 414 lines, 2 matches - Matlab_script/
mvpa_PCC_ambiguity.m , MATLAB, 454 lines, 1 match - Matlab_script/
mvpa_figure.m , MATLAB, 296 lines - Matlab_script/
ppi_config.m , MATLAB, 64 lines - Matlab_script/
run_V1V2_beauty_ppi.m , MATLAB, 309 lines - Matlab_script/
run_all_subjects_step1to , MATLAB, 363 lines4.m - Matlab_script/
run_firstlevel_ambiguity , MATLAB, 231 lines_batch.m - Matlab_script/
run_firstlevel_pm_batch. , MATLAB, 121 linesm - Matlab_script/
run_ppi_firstlevel_batch , MATLAB, 434 lines.m - Matlab_script/
run_ppi_firstlevel_batch , MATLAB, 449 lines, 1 match_manualPPI.m - Matlab_script/
run_secondlevel_ambiguit , MATLAB, 67 linesy_all.m - Matlab_script/
run_secondlevel_beauty_o , MATLAB, 61 linesnesample.m - R_script/
mri_behavior.R , R, 435 lines
The paper's code and data availability statement is in the Data section.
Tracing map
Proposed by the machine: these links were found in the paper and verified at the source, without human review. The map will receive a Zenodo DOI once one of the paper's authors has validated it with their ORCID.
What the map holds:
- 1 repository of the authors' code, each at its verified commit, with its license and how the link was found in the paper;
- 13 scripts, each with its path and the digest of its content;
- 4 matches between paragraphs of the paper and lines of the code (method lexical-v1);
- neither the text of the paper nor the code itself.
Its JSON (tracing-map.json) is deposited on Zenodo with its DOI once the map is validated.
Data
No dataset and no data link were found in the paper.
Data availability statement
Individual-level MRI data are not publicly available due to participant privacy considerations but may be shared upon reasonable request to the corresponding author. Analysis scripts (including the nonparametric SnPM scripts) are available on OSF at https://
Reproduced under the paper's license (CC BY), from the paper cited above.
Versions
The history of this record: each version stored by the harvester or made by a correction of its authors or of the maintainers of its code, and what changed in its facts. The texts of the paper (its abstract, its availability statements) are not part of it; versions that changed only those are not listed.
Version 2, 28 September 2026
- Funding: added Japan Society for the Promotion of Science: 22KJ1813, 23K22374
Version 1, 27 September 2026: the first record
Recorded: type, language, journal, volume, pages, dates, 2 authors, 5 keywords, 47 references.
Cite
This paper
Hitsuwari, J., & Nomura, M. (2026). Neural substrates of ambiguity and beauty in haiku poetry: an fMRI study. Frontiers in human neuroscience, 20, 1887834. https://
BibTeX
@article{hitsuwari2026ne
author = {Hitsuwari, Jimpei and Nomura, Michio},
title = {{Neural substrates of ambiguity and beauty in haiku poetry: an fMRI study}},
journal = {Frontiers in human neuroscience},
year = {2026},
month = aug,
volume = {20},
pages = {1887834},
publisher = {Frontiers Media SA},
issn = {1662-5161},
doi = {10.3389/
url = {https://
pmid = {42729248},
pmcid = {PMC13562011}
}
RIS
TY - JOUR
AU - Hitsuwari, Jimpei
AU - Nomura, Michio
TI - Neural substrates of ambiguity and beauty in haiku poetry: an fMRI study
T2 - Frontiers in human neuroscience
J2 - Front Hum Neurosci
PY - 2026
DA - 2026/
VL - 20
SP - 1887834
SN - 1662-5161
PB - Frontiers Media SA
DO - 10.3389/
UR - https://
LA - en
ER -
CSL-JSON
{
"id": "10.3389/
"type": "article-journal",
"title": "Neural substrates of ambiguity and beauty in haiku poetry: an fMRI study",
"container-title": "Frontiers in human neuroscience",
"author": [
{
"family": "Hitsuwari",
"given": "Jimpei"
},
{
"family": "Nomura",
"given": "Michio"
}
],
"container-title-short":
"volume": "20",
"page": "1887834",
"DOI": "10.3389/
"PMID": "42729248",
"PMCID": "PMC13562011",
"ISSN": "1662-5161",
"publisher": "Frontiers Media SA",
"URL": "https://
"language": "en",
"issued": {
"date-parts": [
[
2026,
8,
28
]
]
}
}
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